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Traveling through Italy - Understanding train transportation

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I will be arriving in Rome and want to get directly onto a high speed train to Florence but am having trouble figuring out which station/train company to use. Also it looks as though I have to have a reservation for the high speed trains? My fear is the plane could be delayed and then what? Also, the best transportation from FCO to the train station and how much time to allow once I land, get through customs and to the station.

Thanks for any help !

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