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International flight, 2 different airlines. Do I have to transfer my bags?

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We are flying American, Boston to Heathrow, then Alitalia ,Heathrow to Rome. Purchased tickets through Orbitz as one ticket. Now I am a little worried as we only have a 2 hour 15 minute layover and have to go from Terminal 5 to 4. Will we have to clear customs again also? If we have a problem and miss our connection how do we book another flight? Through Orbitz or Alitalia or American if it was their delay??/ Thanks for any help.

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